21039559

not annotated - annotated - LINNAEUS only

Systemic virus-induced gene silencing allows functional characterization of maize genes during biotrophic interaction with Ustilago maydis.

Infection of maize (Zea mays) plants with the corn smut fungus Ustilago maydis leads to the formation of large tumors on the stem, leaves and inflorescences. In this biotrophic interaction, plant defense responses are actively suppressed by the pathogen, and previous transcriptome analyses of infected maize plants showed massive and stage-specific changes in host gene expression during disease progression. To identify maize genes that are functionally involved in the interaction with U. maydis, we adapted a virus-induced gene silencing (VIGS) system based on the brome mosaic virus (BMV) for maize. Conditions were established that allowed successful U. maydis infection of BMV-preinfected maize plants. This set-up enabled quantification of VIGS and its impact on U. maydis infection using a quantitative real-time PCR (qRT-PCR)-based readout. In proof-of-principle experiments, an U. maydis-induced terpene synthase was shown to negatively regulate disease development while a protein involved in cell death inhibition was required for full virulence of U. maydis. The results suggest that this system is a versatile tool for the rapid identification of maize genes that determine compatibility with U. maydis.



Ann file

T1	Species 76 81	maize

N1 Reference T1 Taxonomy:4577

T2 Species 123 138 Ustilago maydis

N2 Reference T2 Taxonomy:5270

T3 Species 155 160 maize

N3 Reference T3 Taxonomy:4577

T4 Species 162 170 Zea mays

N4 Reference T4 Taxonomy:4577

T5 Species 188 204 corn smut fungus

T6 Species 205 220 Ustilago maydis

N6 Reference T6 Taxonomy:5270

T7 Species 445 450 maize

N7 Reference T7 Taxonomy:4577

T8 Species 564 569 maize

N8 Reference T8 Taxonomy:4577

T9 Species 631 640 U. maydis

N9 Reference T9 Taxonomy:5270

T10 Species 740 745 maize

N10 Reference T10 Taxonomy:4577

T11 Species 799 808 U. maydis

N11 Reference T11 Taxonomy:5270

T12 Species 913 922 U. maydis

N12 Reference T12 Taxonomy:5270

T13 Species 1031 1040 U. maydis

N13 Reference T13 Taxonomy:5270

T14 Species 1304 1309 maize

N14 Reference T14 Taxonomy:4577

T15 Species 1350 1359 U. maydis

N15 Reference T15 Taxonomy:5270

T16 Species 1204 1213 U. maydis

T17 Species 838 843 maize

N16 Reference T16 Taxonomy:5270 Ustilago maydis

N17 Reference T17 Taxonomy:4577 Zea mays

N5 Reference T5 Taxonomy:5270 Ustilago maydis

#1 AnnotatorNotes T5 TODO check

T18 Species 711 729 brome mosaic virus

N18 Reference T18 Taxonomy:12302 Brome mosaic virus

T19 Species 731 734 BMV

N19 Reference T19 Taxonomy:12302 Brome mosaic virus

T20 Species 822 825 BMV

N20 Reference T20 Taxonomy:12302 Brome mosaic virus