21418208

not annotated - annotated - LINNAEUS only

Soil nutrient status determines how elephant utilize trees and shape environments.

1. Elucidation of the mechanism determining the spatial scale of patch selection by herbivores has been complicated by the way in which resource availability at a specific scale is measured and by vigilance behaviour of the herbivores themselves. To reduce these complications, we studied patch selection by an animal with negligible predation risk, the African elephant. 2. We introduce the concept of nutrient load as the product of patch size, number of patches and local patch nutrient concentration. Nutrient load provides a novel spatially explicit expression of the total available nutrients a herbivore can select from. 3. We hypothesized that elephant would select nutrient-rich patches, based on the nutrient load per 2500 m(2) down to the individual plant scale, and that this selection will depend on the nitrogen and phosphorous contents of plants. 4. We predicted that elephant would cause more adverse impact to trees of lower value to them in order to reach plant parts with higher nutrient concentrations such as bark and root. However, elephant should maintain nutrient-rich trees by inducing coppicing of trees through re-utilization of leaves. 5. Elephant patch selection was measured in a homogenous tree species stand by manipulating the spatial distribution of soil nutrients in a large field experiment using NPK fertilizer. 6. Elephant were able to select nutrient-rich patches and utilized Colophospermum mopane trees inside these patches more than outside, at scales ranging from 2500 down to 100 m(2) . 7. Although both nitrogen and phosphorus contents of leaves from C. mopane trees were higher in fertilized and selected patches, patch choice correlated most strongly with nitrogen content. As predicted, stripping of leaves occurred more in nutrient-rich patches, while adverse impact such as uprooting of trees occurred more in nutrient-poor areas. 8. Our results emphasize the necessity of including scale-dependent selectivity in foraging studies and how elephant foraging behaviour can be used as indicators of change in the availability of nutrients.



Ann file

T1	Out-of-scope 36 44	elephant

T2 Species 439 455 African elephant

N2 Reference T2 Taxonomy:9785

T3 Out-of-scope 737 745 elephant

T4 Out-of-scope 968 976 elephant

T5 Out-of-scope 1139 1147 elephant

T6 Out-of-scope 1252 1260 Elephant

T7 Species 1437 1445 Elephant

N7 Reference T7 Taxonomy:9785

T8 Species 1501 1522 Colophospermum mopane

N8 Reference T8 Taxonomy:162715

T9 Species 1681 1690 C. mopane

N9 Reference T9 Taxonomy:162715

T10 Out-of-scope 2074 2082 elephant

N11 Reference T1 Taxonomy:9780 Elephantidae

N1 Reference T3 Taxonomy:9780 Elephantidae

N3 Reference T4 Taxonomy:9780 Elephantidae

N4 Reference T5 Taxonomy:9780 Elephantidae

N5 Reference T6 Taxonomy:9780 Elephantidae

N6 Reference T10 Taxonomy:9780 Elephantidae